Note: Only playable with a Freeloader or on an American GameCube.
Lloyd Irving and his friend, Genis Sage accompany the chosen, Colette Brunel and her guardians, Raine Sage and Kratos Aurion on her journey of world regeneration. The purpose of the journey is to replenish Sylvarant with mana, a necessary energy to their survival. The path of the journey consists of five temples which must be unsealed by Colette. On their journey, they meet Sheena Fujibayashi who comes from Tethe'alla, a world parallel to Sylvarant. She tells them about how the two worlds vie for each other's mana through the journey of regeneration. Hoping to find answers from the angel Remiel, the party meets him at the final seal and are betrayed by Remiel and Kratos; both are revealed to be part of Cruxis, an evil organization led by Mithos Yggdrasill.
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